Teleostei (teleosts) >
Gobiiformes (Gobies) >
Gobiidae (Gobies) > Gobionellinae
Etymology: Knipowitschia: Because of N.M.Knipowitsch, a zoologist and Russian ichthyologist from the Academy of Sciences. Expeditions in "Pomor" and "Pervosvanniy" ships to Azov and Black sea (1898-1908).
Eponymy: Nikolai Mikhailovich Knipovich (also given as Knipowitsch) (1862–1939) was a Russian oceanographer, marine zoologist and ichthyologist. [...] (Ref. 128868), visit book page.
Environment: milieu / climate zone / depth range / distribution range
Ecology
Marine; freshwater; brackish; demersal. Temperate; 4°C - 20°C (Ref. 2059); 48°N - 42°N, 30°E - 54°E
Eurasia: Brackish portions of the Black Sea, Sea of Azov and Caspian Sea.
Length at first maturity / Size / Weight / Age
Maturity: Lm 2.0 range ? - ? cm
Max length : 5.0 cm TL male/unsexed; (Ref. 2058); 3.5 cm TL (female)
Dorsal spines (total): 6 - 8; Dorsal soft rays (total): 7 - 9; Anal spines: 1; Anal soft rays: 8 - 9. This species is distinguished from its congeners by the following characters: upper part of caudal fin longer than the lower part; anterior extremity of anterior oculoscapular canal in front of middle of eye; no posterior oculoscapular canal; back unscaled in front of origin of second dorsal; breeding males with 4-8 dark bars when alive; total scales in midlateral series 33-40 (Ref. 59043).
Occur in shallow water, on sand and among weeds (Ref. 4696). Live less than 2 years. Feed on small benthic invertebrates (Ref. 59043). Spawn after first winter. Females may spawn several times during a season. Males defend eggs in cavities under stones, plant material or molluscs shells (Ref. 59043) like underside of lamellibranch valves (Cardium, Monodonta, Unio, etc.) (Ref. 4696). Eggs hatch at 3.0-3.8 mm (Ref. 59043).
Males defend eggs in cavities under stones, plant material or molluscs shells (Ref. 59043). Females spawn several times during a season (Ref. 59043).
Miller, P.J., 1986. Gobiidae. p. 1019-1085. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. Volume 3. UNESCO, Paris. (Ref. 4696)
IUCN Red List Status (Ref. 130435: Version 2024-2)
Threat to humans
Harmless
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Estimates based on models
Phylogenetic diversity index (Ref.
82804): PD
50 = 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00977 (0.00444 - 0.02153), b=3.04 (2.85 - 3.23), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref.
93245).
Trophic level (Ref.
69278): 3.4 ±0.4 se; based on size and trophs of closest relatives
Generation time: 1.7 ( na - na) years. Estimated as median ln(3)/K based on 2
growth studies.
Resilience (Ref.
120179): High, minimum population doubling time less than 15 months (tmax < 2; tm < 1 (8-10 months);).
Fishing Vulnerability (Ref.
59153): Low vulnerability (10 of 100).
