Dorsal-fin rays 9-10; anal-fin rays 8; pectoral-fin rays 17-24; principal caudal-fin rays15, gills3, the third arch with 3 gill rakers; total vertebrae 34-35, with asymmetrical lateral bending starting behind the skull, bent at an angle of 85°-92°; asymmetrical skull in frontal view; naked skin, surface of head and body without striae; disc without adhesive papillae; posterior margin may be partially fringed; region B small, left- and right-hand sides separate, larger on the right-hand side than on left-hand side (Ref. 117884).