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| Sound produced | yes, active sound production |
| Type of sound produced | knocks (very weak) |
| Sound production organ | swim bladder (possibly opercula) |
| Sound mechanism | vibration by contraction of extrinsic muscles associated with swim bladder lobes, reinforced by general contraction of skeletal muscles or violent closing of opercula |
| Behavioural context | under duress (handling or moderate irritaion) |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark | recorded with highpass filter: 30 Hz |