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| Sound produced | yes, active sound production |
| Type of sound produced | grunts, knocks |
| Sound production organ | pneumatic duct & swim bladder |
| Sound mechanism | not specified, but probably vibration of swim bladder and pneumatic duct |
| Behavioural context | when alarmed, (sound was recorded when enclosure lid was lifted) |
| Reference | Tiepelt, H., 2005 |
| Remark | outdoor recording (enclosure submerged in freshwater lake). Spontaneous sound production from 1 specimen after removing the top-cover of enclosure. Sound recorded in presence of 11 others (20-25 cm long) |