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| Sound produced | yes, active sound production |
| Type of sound produced | grunts |
| Sound production organ | pharyngeal teeth & swim bladder |
| Sound mechanism | pharyngeal teeth stridulation amplified by adjacent swim bladder |
| Behavioural context | weak sounds during competitve feeding, louder under duress (netted, handled or with electrical stimulation) |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark | recording amplified by 3 dB, low pass nois reduction filter (< 3600 Hz) applied to recording |