| Key to native species of the Biobio with emphasis on high altitude regions. |
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| Ruiz, V.H. and T.M. Berra 1994. Fishes of the high Biobio River of south-central Chile with notes on diet and speculations on the origin of the ichthyofauna. Ichthyol. Explor. Freshwat. 5(1):5-18. Ref No [26616] Key No. [1164] |
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