Africa: lower and middle Zambezi, Buzi, Pungwe (Ref. 52193), lower Sabi and Ludi Rivers (Ref. 42505), Luapula-Moero-Bangwelo (Zambian Congo system) (Ref. 42505, 52193). Also known from the Ruvuma and Rufiji rivers in Tanzania (Ref. 7248, 52193), lakes Malawi, Tanganyika (Ref. 4967, 52193) and Rukwa, and other eastward flowing rivers in Tanzania (Ref. 4967).