List of Nominal Species of Cottidae
(Sculpins)
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Nominal Species (n = 415) Present allocation (n = 293) Ref.
Oligocottus acuticeps Gilbert, 1896 Clinocottus acuticeps 26282
Cottus aenaeus Mitchill, 1814 Myoxocephalus aenaeus 12965
Cottus affinis Heckel, 1837 Cottus gobio 13696
Crossias albomaculatus Schmidt, 1916 Porocottus mednius 43939
Centridermichthys alcicornis Herzenstein, 1890 Alcichthys elongatus 58496
Cottus aleuticus Gilbert, 1896 Cottus aleuticus 3814
Crossias allisi Jordan & Starks, 1904 Porocottus allisi 26282
Cottus amblystomopsis Schmidt, 1904 Cottus amblystomopsis 559
Oligocottus analis Girard, 1858 Clinocottus analis 12965
Thyriscus anoplus Gilbert & Burke, 1912 Thyriscus anoplus 6876
Artediellus antilope Schmidt, 1937 Artediellina antilope 46206
Artediellus aporosus Soldatov, 1922 Artediellus aporosus 46206
Centridermichthys argenteus Döderlein, 1887 Pseudoblennius argenteus 40966
Ochotskia armata Schmidt, 1916 Icelus armatus 46206
Leptocottus armatus Girard, 1854 Leptocottus armatus 26282
Cottus asper Richardson, 1836 Cottus asper 5723
Cottus asperrimus Rutter, 1908 Cottus asperrimus 5723
Radulinus asprellus Gilbert, 1890 Radulinus asprellus 26282
Artediellus atlanticus Jordan & Evermann, 1898 Artediellus atlanticus 4698
Cottus aturi Freyhof, Kottelat & Nolte, 2005 Cottus aturi 55856
Artedielloides auriculatus Soldatov, 1922 Artedielloides auriculatus 27683
Boreocottus axillaris Gill, 1859 Myoxocephalus scorpioides 50838
Cottus baicalensis Dybowski, 1874 Batrachocottus baicalensis 33021
Callionymus baikalensis Pallas, 1776 Comephorus baikalensis 1830
Cottus baileyi Robins, 1961 Cottus baileyi 5723
Cottus bairdii Girard, 1850 Cottus bairdii 1998
Cottus bathybius Günther, 1878 Zesticelus bathybius 40966
Triglops beani Gilbert, 1896 Triglops pingelii 7297
Cottus beldingii Eigenmann & Eigenmann, 1891 Cottus beldingii 12965
Potamocottus bendirei Bean, 1881 Cottus bendirei 50838
Limnocottus bergianus Taliev, 1935 Limnocottus bergianus 26334
Taurocottus bergii Soldatov & Pavlenko, 1915 Taurocottus bergii 46206
Stelgistrum beringianum Gilbert & Burke, 1912 Stelgistrum beringianum 6876
Cottus bicornis Reinhardt, 1840 Icelus bicornis 26282
Archaulus biseriatus Gilbert & Burke, 1912 Archistes biseriatus 43939
Aspicottus bison Girard, 1854 Enophrys bison 26282
Cottus boleoides Girard, 1850 Cottus cognatus 54621
Radulinus boleoides Gilbert, 1898 Radulinus boleoides 26282
Icelinus borealis Gilbert, 1896 Icelinus borealis 26282
Abyssocottus boulengeri Berg, 1906 Cottinella boulengeri 26282
Porocottus bradfordi Rutter, 1898 Porocottus mednius 43939
Cottus brandtii Steindachner, 1867 Myoxocephalus brandtii 26282
Trichocottus brashnikovi Soldatov & Pavlenko, 1915 Trichocottus brashnikovi 43939
Cottus bubalis Euphrasen, 1786 Taurulus bubalis 3397
Icelinus burchami Evermann & Goldsborough, 1907 Icelinus burchami 26282
Cottus caeruleomentum Kinziger, Raesly & Neely, 2000 Cottus caeruleomentum 37399
Artediellus camchaticus Gilbert & Burke, 1912 Artediellus camchaticus 559
Crossias camtschaticus Schmidt, 1916 Porocottus camtschaticus 33021
Icelus canaliculatus Gilbert, 1896 Icelus canaliculatus 6793
Potamocottus carolinae Gill, 1861 Cottus carolinae 12965
Agonocottus cataphractus Pavlenko, 1910 Icelus cataphractus 26282
Sigmistes caulias Rutter, 1898 Sigmistes caulias 26282
Icelinus cavifrons Gilbert, 1890 Icelinus cavifrons 2850
Podabrus centropomus Richardson, 1848 Vellitor centropomus 26282
Cottus chamberlaini Evermann & Goldsborough, 1907 Cottus cognatus 1998
Cottus chattahoochee Neely, Williams & Mayden, 2007 Cottus chattahoochee 72414
Blennicottus clarki Evermann & Goldsborough, 1907 Artediellus pacificus 11366
Cottus claviger Cuvier, 1829 Enophrys diceraus 50838
Cottus cognatus Richardson, 1836 Cottus cognatus 10280
Cottus comephoroides Berg, 1900 Cottocomephorus inermis 12480
Stelgistrum concinnum Andriashev, 1935 Stelgistrum concinnum 43939
Cottus confusus Bailey & Bond, 1963 Cottus confusus 5723
Allartedius corallinus Hubbs, 1926 Artedius corallinus 12965
Porocottus coronatus Yabe, 1992 Porocottus coronatus 34715
Podabrus cottoides Richardson, 1848 Pseudoblennius cottoides 26282
Ruscariops creaseri Hubbs, 1926 Ruscarius creaseri 12965
Phasmatocottus ctenopterygius Bolin, 1936 Phasmatocottus ctenopterygius 559
Cottus cyclophthalmus Sideleva, Kesminas & Zhidkov, 2022 Cottus cyclophthalmus 129290
Cottus czerskii Berg, 1913 Cottus czerskii 26334
Cottus decastrensis Kner, 1865 Myoxocephalus stelleri 58496
Artedius delacyi Hubbs & Schultz, 1941 Artedius lateralis 10937
Radulinopsis derjavini Soldatov & Lindberg, 1930 Radulinopsis derjavini 45347
Radulinopsis derjugini Soldatov, 1930 Radulinopsis derjavini 45347
Gymnocanthus detrisus Gilbert & Burke, 1912 Gymnocanthus detrisus 559
Cottus diceraus Pallas, 1787 Enophrys diceraus 58462
Stlengis distoechus Bolin, 1936 Stlengis distoechus 559
Triglops dorothy Pietsch & Orr, 2006 Triglops dorothy 57076
Cottus duranii Freyhof, Kottelat & Nolte, 2005 Cottus duranii 55856
Comephorus dybowskii Korotneff, 1904 Comephorus dybowskii 12480
Artediellus dydymovi Soldatov, 1915 Artediellus dydymovi 26282
Cottus dzungaricus Kottelat, 2006 Cottus dzungaricus 57778
Cottus echinatus Bailey & Bond, 1963 Cottus echinatus 3814
Icelus ecornis Tsutsui & Yabe, 1996 Icelus ecornis 41299
Antipodocottus elegans Fricke & Brunken, 1984 Antipodocottus elegans 7300
Centridermichthys elegans Steindachner, 1881 Bero elegans 26282
Abyssocottus elochini Taliev, 1955 Abyssocottus elochini 26334
Centridermichthys elongatus Steindachner, 1881 Alcichthys elongatus 46206
Oligocottus embryum Jordan & Starks, 1895 Clinocottus embryum 26282
Ainocottus ensiger Jordan & Starks, 1904 Myoxocephalus polyacanthocephalus 10937
Artediellus europaeus Knipowitsch, 1907 Artediellus atlanticus 93826
Icelus euryops Bean, 1890 Icelus euryops 26282
Bolinia euryptera Yabe, 1991 Bolinia euryptera 39612
Cottus extensus Bailey & Bond, 1963 Cottus extensus 5723
Trachidermus fasciatus Heckel, 1837 Trachidermus fasciatus 26763
Artedius fenestralis Jordan & Gilbert, 1883 Artedius fenestralis 26282
Cottus ferrugineus Bonaparte, 1846 Cottus gobio 55856
Cottus ferrugineus Heckel & Kner, 1858 Cottus gobio 55856
Icelinus filamentosus Gilbert, 1890 Icelinus filamentosus 6793
Icelinus fimbriatus Gilbert, 1890 Icelinus fimbriatus 2850
Elanura forficata Gilbert, 1896 Triglops forficatus 7297
Cottus formosus Girard, 1851 Cottus cognatus 1998
Cottus franklini Agassiz, 1850 Cottus cognatus 1998
Artediellus fuscimentus Nelson, 1986 Artediellus fuscimentus 41299
Abyssocottus fuscus Bogdanov, 2014 Abyssocottus fuscus 130169
Antipodocottus galatheae Bolin, 1952 Antipodocottus galatheae 7300
Gymnocanthus galeatus Bean, 1881 Gymnocanthus galeatus 6793
Abyssocottus gibbosus Berg, 1906 Abyssocottus gibbosus 26334
Abyssocottus gibbosus subulatus Dybowski, 1908 Abyssocottus subulatus 130169
Hemilepidotus gilberti Jordan & Starks, 1904 Hemilepidotus gilberti 559
Icelus gilberti Taranetz, 1936 Icelus gilberti 559
Synchirus gilli Bean, 1890 Synchirus gilli 26282
Cottus girardi (non Robins, 1961) Cottus bairdii 1998
Cottus girardi Robins, 1961 Cottus girardi 5723
Oligocottus globiceps Girard, 1858 Clinocottus globiceps 26282
Cottus gobio Linnaeus, 1758 Cottus gobio 55856
Cottus gobio haemusi (non Marinov & Dikov, 1986) Cottus gobio 59043
Cottus gobio haemusi Marinov & Dikov, 1986 Cottus haemusi 59043
Cottus gobio hispaniolensis (non Băcescu & Băcescu-Meşter, 1964) Cottus gobio 55856
Cottus gobio hispaniolensis Băcescu & Băcescu-Meşter, 1964 Cottus hispaniolensis 55856
Cottus gobio jaxartensis Berg, 1916 Cottus gobio 13696
Cottus gobio macrostomus Jeitteles, 1863 Cottus gobio 55856
Cottus gobio microcephalus (non Kessler, 1868) Alpinocottus poecilopus 55856
Cottus gobio microcephalus Kessler, 1868 Cottus koshewnikowi 55856
Cottus gobio milvensis (non Soldatov, 1924) Cottus gobio 55856
Cottus gobio milvensis Soldatov, 1924 Cottus koshewnikowi 55856
Cottus gobio pellegrini Băcescu & Băcescu-Meşter, 1964 Cottus gobio 13696
Cottus gobio pellegrini Vladykov, 1931 Cottus gobio 13696
Cottus gobio roseus Odenwall, 1927 Cottus gobio 13696
Cottus godlewskii Dybowski, 1874 Limnocottus godlewskii 33021
Limnocottus godlewskii bergi Dybowski, 1908 Limnocottus godlewskii 2058
Abyssocottus godlewskii griseus Taliev, 1955 Limnocottus griseus 40966
Artediellus gomojunovi Taranetz, 1933 Artediellus gomojunovi 6876
Cottiusculus gonez Jordan & Starks, 1904 Cottiusculus gonez 559
Procottus gotoi Sideleva, 2001 Procottus gotoi 53210
Cottus gracilis Heckel, 1837 Cottus cognatus 1998
Cottus gratzianowi Sideleva, Naseka & Zhidkov, 2015 Cottus gratzianowi 101366
Uranidea greenei Gilbert & Culver, 1898 Cottus greenei 33021
Cottocomephorus grewingki alexandrae Taliev, 1935 Cottocomephorus alexandrae 51998
Cottus grewingkii Dybowski, 1874 Cottocomephorus grewingkii 2058
Cottus groenlandicus Cuvier, 1829 Myoxocephalus scorpius 43939
Cottopsis gulosus Girard, 1854 Cottus gulosus 26282
Metacottus gurwicii Taliev, 1946 Procottus gurwicii 26282
Cottus haitej Dybowski, 1869 Mesocottus haitej 26282
Cottus hangiongensis Mori, 1930 Cottus hangiongensis 559
Axyrias harringtoni Starks, 1896 Artedius harringtoni 26282
Cottus hemilepidotus Tilesius, 1811 Hemilepidotus hemilepidotus 26282
Asprocottus herzensteini Berg, 1906 Asprocottus herzensteini 130169
Gymnocanthus herzensteini Jordan & Starks, 1904 Gymnocanthus herzensteini 559
Asprocottus herzensteini abyssalis Taliev, 1955 Asprocottus abyssalis 130169
Asprocottus herzensteini intermedius Taliev, 1955 Asprocottus intermedius 46206
Asprocottus herzensteini parmiferus Taliev, 1955 Asprocottus parmiferus 26282
Asprocottus herzensteini platycephalus Taliev, 1955 Asprocottus platycephalus 26282
Cottus hexacornis Richardson, 1823 Myoxocephalus quadricornis 26282
Leiocottus hirundo Girard, 1856 Leiocottus hirundo 2850
Uranidea hoyi Putnam, 1876 Cottus cognatus 1998
Cottus hubbsi (non Bailey & Dimick, 1949) Cottus bairdii 1998
Cottus hubbsi Bailey & Dimick, 1949 Cottus hubbsi 54825
Cottus humilis Bean, 1881 Myoxocephalus jaok 43939
Cottus hypselurus Robins & Robison, 1985 Cottus hypselurus 3814
Pegedictis ictalops Rafinesque, 1820 Cottus bairdii 93641
Cottus immaculatus Kinziger & Wood, 2010 Cottus immaculatus 83416
Myoxocephalus incitus Watanabe, 1958 Myoxocephalus stelleri 43239
Cottus inermis Yakovlev, 1890 Cottocomephorus inermis 12480
Artediellus ingens Nelson, 1986 Artediellus ingens 43939
Cottus intermedius Temminck & Schlegel, 1843 Gymnocanthus intermedius 26282
Furcina ishikawae Jordan & Starks, 1904 Furcina ishikawae 559
Cottus jaok Cuvier, 1829 Myoxocephalus jaok 26282
Icelinus japonicus Yabe, Tsumura & Katayama, 1980 Icelinus japonicus 559
Porocottus japonicus Schmidt, 1935 Porocottus japonicus 34715
Procottus jeittelesi bicolor Dybowski, 1908 Procottus jeittelesii 2058
Procottus jeittelesi major Taliev, 1949 Procottus major 33021
Procottus jeittelesi minor Taliev, 1946 Procottus gurwicii 46206
Cottus jeittelesii Dybowski, 1874 Procottus jeittelesii 26282
Elanura jordani Schmidt, 1903 Triglops jordani 7297
Hemilepidotus jordani Bean, 1881 Hemilepidotus jordani 6793
Prionistius jordani Jordan & Starks, 1904 Triglops jordani 54621
Cottus kaganowskii Berg, 1932 Cottus cognatus 54621
Cottus kanawhae Robins, 2005 Cottus kanawhae 54473
Cottus kazika Jordan & Starks, 1904 Cottus kazika 559
Paracottus kessleri arachlensis Tarchova, 1962 Leocottus kesslerii 57778
Paracottus kessleri gussinensis Tarchova, 1962 Leocottus kesslerii 57778
Cottus kesslerii Dybowski, 1874 Leocottus kesslerii 26334
Cottus kesslerii bauntovi Taliev, 1946 Leocottus kesslerii 50838
Cottus klamathensis Gilbert, 1898 Cottus klamathensis 5723
Cottus klamathensis polyporus Daniels & Moyle, 1984 Cottus klamathensis 54621
Paracottus kneri putorania Koryakov & Sidelov, 1976 Paracottus knerii 50600
Cottus knerii Dybowski, 1874 Paracottus knerii 26282
Cottus kolymensis Sideleva & Goto, 2012 Cottus kolymensis 93142
Cottus koreanus Fujii, Choi & Yabe, 2005 Cottus koreanus 93142
Asprocottus korjakovi Sideleva, 2001 Asprocottus korjakovi 53210
Asprocottus korjakovi minor Sideleva, 2001 Asprocottus minor 53210
Abyssocottus korotneffi Berg, 1906 Abyssocottus korotneffi 42042
Cottus koshewnikowi (non Gratzianov, 1907) Cottus gobio 55856
Cottus koshewnikowi Gratzianov, 1907 Cottus koshewnikowi 55856
Limnocottus kozovi Taliev, 1946 Cyphocottus megalops 46206
Uranidea kumlieni Hoy, 1876 Cottus bairdii 54621
Cottus kuznetzovi (non Berg, 1903) Alpinocottus poecilopus 93142
Cottus kuznetzovi Berg, 1903 Cottus kuznetzovi 93142
Scorpaenichthys lateralis Girard, 1854 Artedius lateralis 26282
Acanthocottus laticeps Gilbert, 1896 Megalocottus platycephalus 26282
Cottus latifrons Gratzianov, 1904 Myoxocephalus quadricornis 13696
Stelgidonotus latifrons Gilbert & Thompson, 1905 Oligocottus rimensis 43939
Cottus leiopomus Gilbert & Evermann, 1894 Cottus leiopomus 5723
Astrocottus leprops Bolin, 1936 Astrocottus leprops 76844
Astrocottus leprops (non Bolin, 1936) Astrocottus matsubarae 76844
Porocottus leptosomus Muto, Choi & Yabe, 2002 Porocottus leptosomus 44254
Cottus lilljeborgii Collett, 1875 Micrenophrys lilljeborgii 12965
Icelinus limbaughi Rosenblatt & Smith, 2004 Icelinus limbaughi 75898
Ceratocottus lucasi (non Jordan & Gilbert, 1898) Enophrys diceraus 11366
Ceratocottus lucasi Jordan & Gilbert, 1898 Enophrys lucasi 33021
Prionistius macellus Bean, 1884 Triglops macellus 7297
Pterygiocottus macouni Bean & Weed, 1920 Artedius harringtoni 4925
Cottus macrops Rutter, 1908 Cottus klamathensis 50838
Cottus maculatus Fischer, 1885 Taurulus bubalis 9981
Oligocottus maculosus Girard, 1856 Oligocottus maculosus 26282
Icelus mandibularis Yabe, 1983 Icelus mandibularis 559
Uranidea marginata Bean, 1881 Cottus marginatus 12965
Centridermichthys marmoratus Döderlein, 1884 Pseudoblennius marmoratus 26282
Cottus marmoratus Valenciennes, 1832 Myoxocephalus stelleri 58496
Ocynectes maschalis Jordan & Starks, 1904 Ocynectes maschalis 559
Astrocottus matsubarae Katayama, 1942 Astrocottus matsubarae 76844
Myoxocephalus matsubarai Watanabe, 1958 Myoxocephalus matsubarai 559
Microcottus matuaensis Yabe & Pietsch, 2003 Microcottus matuaensis 49856
Ruscarius meanyi Jordan & Starks, 1895 Ruscarius meanyi 3814
Myoxocephalus mednius Bean, 1898 Porocottus mednius 43939
Andriashevicottus megacephalus Fedorov, 1990 Andriashevicottus megacephalus 41742
Antipodocottus megalops DeWitt, 1969 Antipodocottus megalops 5755
Cottus megalops Gratzianov, 1902 Cyphocottus megalops 26282
Asprocottus megalops eurystomus Taliev, 1955 Cyphocottus eurystomus 26282
Cottus meridionalis Girard, 1850 Cottus bairdii 37399
Stlengis mesembrinus Fricke & Brunken, 1983 Antipodocottus mesembrinus 34698
Cottus metae Freyhof, Kottelat & Nolte, 2005 Cottus metae 55856
Triglops metopias Gilbert & Burke, 1912 Triglops metopias 7297
Artediellus miacanthus Gilbert & Burke, 1912 Artediellus miacanthus 6876
Uranidea microstoma Lockington, 1880 Cottus aleuticus 1998
Cottus microstomus (non Heckel, 1837) Cottus gobio 55856
Cottus microstomus Heckel, 1837 Cottus microstomus 55856
Cottiusculus minor Watanabe, 1958 Artediellus minor 40966
Cottus minutus Pallas, 1814 Porocottus minutus 33021
Vellitor minutus Iwata, 1983 Vellitor minutus 559
Cottus minutus volki Taranetz, 1933 Alpinocottus volki 26282
Schmidtia misakia Jordan & Starks, 1904 Stlengis misakia 26282
Ocynectes modestus Snyder, 1911 Ocynectes modestus 34595
Triglops murrayi Günther, 1888 Triglops murrayi 7297
Cottus nasalis Berg, 1933 Cottus nasalis 26334
Artediellus neyelovi Muto, Yabe & Amaoka, 1994 Artediellus neyelovi 41299
Cottus niger Bean, 1881 Myoxocephalus niger 26282
Artediellus nigripinnis Schmidt, 1937 Artediellichthys nigripinnis 43939
Cottiusculus nihonkaiensis Kai & Nakabo, 2009 Cottiusculus nihonkaiensis 83439
Cottus nikolskii Berg, 1900 Batrachocottus nikolskii 26282
Batrachocottus nikolskii multiradiatus Berg, 1907 Batrachocottus multiradiatus 26282
Artedius notospilotus Girard, 1856 Artedius notospilotus 2850
Cottus nozawae Snyder, 1911 Cottus nozawae 559
Ricuzenius nudithorax Bolin, 1936 Ricuzenius nudithorax 559
Triglops nybelini Jensen, 1944 Triglops nybelini 7297
Phallocottus obtusus Schultz, 1938 Phallocottus obtusus 2850
Artediellus ochotensis Gilbert & Burke, 1912 Artediellus ochotensis 559
Icelus ochotensis Schmidt, 1927 Icelus ochotensis 559
Zesticelus ochotensis Yabe, 1995 Zesticelus ochotensis 13470
Cottus octodecemspinosus Mitchill, 1814 Myoxocephalus octodecemspinosus 12965
Icelinus oculatus (non Gilbert, 1890) Icelinus fimbriatus 11980
Icelinus oculatus Gilbert, 1890 Icelinus oculatus 3814
Alcichthys okiensis Mori, 1956 Alcichthys elongatus 41299
Triglops ommatistius Gilbert, 1913 Triglops murrayi 7297
Triglops ommatistius terraenovae Gilbert, 1913 Triglops murrayi 7297
Cottus onychus Eigenmann & Eigenmann, 1892 Cottus ricei 1998
Stlengis osensis Jordan & Starks, 1904 Stlengis osensis 559
Furcina osimae Jordan & Starks, 1904 Furcina osimae 559
Astrocottus oyamai Watanabe, 1958 Astrocottus oyamai 559
Artediellus pacificus Gilbert, 1896 Artediellus pacificus 26282
Limnocottus pallidus Taliev, 1948 Limnocottus pallidus 26334
Melletes papilio Bean, 1880 Hemilepidotus papilio 26282
Myoxocephalus parvulus Gilbert & Burke, 1912 Microcottus sellaris 49856
Cottopsis parvus Girard, 1854 Cottus asper 1998
Cottus paulus Williams, 2000 Cottus paulus 36875
Paracottus pelagicus Taliev, 1955 Leocottus kesslerii 85815
Pseudoblennius percoides Günther, 1861 Pseudoblennius percoides 559
Cottus perifretum Freyhof, Kottelat & Nolte, 2005 Cottus perifretum 55856
Icelus perminovi Taranetz, 1936 Icelus perminovi 559
Cottus perplexus Gilbert & Evermann, 1894 Cottus perplexus 5723
Cottus petiti Băcescu & Băcescu-Meşter, 1964 Cottus petiti 13696
Cottus philonips Eigenmann & Eigenmann, 1892 Cottus cognatus 1998
Icelinus pietschi Yabe, Soma & Amaoka, 2001 Icelinus pietschi 45337
Ricuzenius pinetorum Jordan & Starks, 1904 Ricuzenius pinetorum 559
Triglops pingelii Reinhardt, 1837 Triglops pingelii 12965
Triglops pingelii islandicus Jensen, 1944 Triglops murrayi 7297
Triglops pingelii pacificus Schmidt, 1930 Triglops pingelii 10937
Triglops pingelii pietschmanni Jensen, 1944 Triglops murrayi 7297
Triglops pingelii typicus Le Danois, 1914 Triglops pingelii 7297
Triglops pingelii var. subborealis Le Danois, 1914 Triglops murrayi 7297
Cottus pistilliger Pallas, 1814 Gymnocanthus pistilliger 26282
Cottus pitensis Bailey & Bond, 1963 Cottus pitensis 5723
Cottus platycephalus Pallas, 1814 Megalocottus platycephalus 12965
Triglops pleurostictus Cope, 1865 Triglops pingelii 7297
Archistes plumarius Jordan & Gilbert, 1898 Archistes plumarius 43939
Cottus poecilopus Heckel, 1837 Alpinocottus poecilopus 55856
Cottus poecilopus altaicus (non Kaschenko, 1899) Alpinocottus poecilopus 57778
Cottus poecilopus altaicus Kaschenko, 1899 Cottus altaicus 57778
Cottus poecilopus macrostomus Jeitteles, 1862 Alpinocottus poecilopus 13696
Cottus poecilopus microstomus Jeitteles, 1862 Alpinocottus poecilopus 13696
Uranidea pollicaris Jordan & Gilbert, 1882 Cottus ricei 1998
Cottus pollux Günther, 1873 Cottus pollux 559
Cottus polyacanthocephalus Pallas, 1814 Myoxocephalus polyacanthocephalus 26282
Cottiusculus primoricus Prokofiev, 2020 Cottiusculus primoricus 123154
Cottus princeps Gilbert, 1898 Cottus princeps 5723
Acanthocottus profundorum Gilbert, 1896 Zesticelus profundorum 26282
Cottus protrusus Schultz & Spoor, 1933 Cottus aleuticus 10280
Artedius pugetensis Steindachner, 1876 Chitonotus pugetensis 26282
Asprocottus pulcher Taliev, 1955 Asprocottus pulcher 26334
Abyssocottus pumilus Bogdanov, 2014 Abyssocottus pumilus 130169
Potamocottus punctulatus Gill, 1861 Cottus bairdii 54621
Cottus pygmaeus Williams, 1968 Cottus paulus 5723
Porocottus quadratus Bean, 1898 Microcottus sellaris 49856
Cottus quadricornis Linnaeus, 1758 Myoxocephalus quadricornis 86838
Cottus quadricornis asundensis Lönnberg, 1932 Myoxocephalus quadricornis 13696
Cottus quadricornis borkensis Lönnberg, 1939 Myoxocephalus quadricornis 13696
Cottus quadricornis frykenensis Lönnberg, 1932 Myoxocephalus quadricornis 13696
Cottus quadricornis kallavesensis Lönnberg, 1933 Myoxocephalus quadricornis 13696
Cottus quadricornis lonnbergi Lönnberg, 1919 Myoxocephalus quadricornis 13696
Myoxocephalus quadricornis lonnbergi Berg, 1916 Myoxocephalus quadricornis 13696
Cottus quadricornis oernensis Lönnberg, 1932 Myoxocephalus quadricornis 13696
Myoxocephalus quadricornis onegensis Berg & Popov, 1932 Myoxocephalus quadricornis 13696
Cottus quadricornis pygmaeus Lönnberg, 1933 Myoxocephalus quadricornis 13696
Cottus quadricornis relicta Lilljeborg, 1891 Myoxocephalus quadricornis 13696
Cottus quadricornis vaenernensis Lönnberg, 1932 Myoxocephalus quadricornis 13696
Cottus quadricornis vermelnensis Nybelin, 1940 Myoxocephalus quadricornis 13696
Porocottus quadrifilis Gill, 1859 Porocottus quadrifilis 6876
Artedius quadriseriatus Lockington, 1880 Icelinus quadriseriatus 12965
Uranidea quiescens DeKay, 1842 Cottus cognatus 1998
Myoxocephalus raninus Jordan & Starks, 1904 Myoxocephalus stelleri 43939
Icelus rastrinoides Taranetz, 1936 Icelus rastrinoides 27683
Blennicottus recalvus Greeley, 1899 Clinocottus recalvus 12965
Astrocottus regulus Tsuruoka, Maruyama & Yabe, 2008 Astrocottus regulus 76844
Cottus reinii Hilgendorf, 1879 Cottus reinii 559
Cottus rhenanus Freyhof, Kottelat & Nolte, 2005 Cottus rhenanus 55856
Ascelichthys rhodorus Jordan & Gilbert, 1880 Ascelichthys rhodorus 26282
Uranidea rhothea Smith, 1882 Cottus rhotheus 1998
Cottopsis ricei Nelson, 1876 Cottus ricei 1998
Cottus richardsoni Agassiz, 1850 Cottus bairdii 1998
Trachidermus richardsonii Heckel, 1837 Cottus asper 1998
Rusciculus rimensis Greeley, 1899 Oligocottus rimensis 26282
Cottus rondeleti Freyhof, Kottelat & Nolte, 2005 Cottus rondeleti 55856
Eximia rubellio Greeley, 1899 Oligocottus rubellio 12965
Cottus sabaudicus Sideleva, 2009 Cottus sabaudicus 81212
Daruma sagamia Jordan & Starks, 1904 Daruma sagamia 559
Artediellus scaber Knipowitsch, 1907 Artediellus scaber 4698
Artediellus scaber beringianus Schmidt, 1927 Artediellus scaber 10937
Cottus scaturigo Freyhof, Kottelat & Nolte, 2005 Cottus gobio 129293
Triglops scepticus Gilbert, 1896 Triglops scepticus 7297
Cottus schitsuumsh Lemoine, Young, Mckelvey, Eby, Pilgrim & Schwartz, 2014 Cottus schitsuumsh 94998
Artediellus schmidti Soldatov, 1915 Artediellus schmidti 26282
Cottiusculus schmidti (non Jordan & Starks, 1904) Cottiusculus nihonkaiensis 83439
Cottiusculus schmidti Jordan & Starks, 1904 Cottiusculus schmidti 559
Cottus scorpioides Fabricius, 1780 Myoxocephalus scorpioides 80565
Cottus scorpius Linnaeus, 1758 Myoxocephalus scorpius 43939
Icelus scutiger Bean, 1890 Rastrinus scutiger 6876
Icelus sekii Tsuruoka, Munehara & Yabe, 2006 Icelus sekii 58328
Acanthocottus sellaris Gilbert, 1896 Microcottus sellaris 26282
Porocottus sellaris ochotensis Schmidt, 1929 Microcottus sellaris 49856
Cottopsis semiscaber Cope, 1872 Cottus bairdii 93641
Cottus sibiricus Warpachowski, 1889 Cottus sibiricus 26334
Cottus sibiricus altaicus Li & Ho, 1966 Cottus dzungaricus 57778
Cottus sinensis Sauvage, 1873 Myoxocephalus sinensis 46206
Lepidobero sinensis Qin & Jin, 1992 Lepidobero sinensis 31517
Sigmistes smithi Schultz, 1938 Sigmistes smithi 45337
Oligocottus snyderi Greeley, 1898 Oligocottus snyderi 26282
Icelus spatula Gilbert & Burke, 1912 Icelus spatula 7251
Cottus specus Adams & Burr, 2013 Cottus specus 92978
Icelus spiniger Gilbert, 1896 Icelus spiniger 6793
Hemilepidotus spinosus Ayres, 1854 Hemilepidotus spinosus 12965
Cottus spinulosus Kessler, 1872 Cottus spinulosus 2058
Stelgistrum stejnegeri Jordan & Gilbert, 1898 Stelgistrum stejnegeri 559
Myoxocephalus stelleri Tilesius, 1811 Myoxocephalus stelleri 43939
Cottus szanaga Dybowski, 1869 Cottus szanaga 53210
Cottus taeniopterus (non Kner, 1868) Megalocottus platycephalus 11366
Cottus taeniopterus Kner, 1868 Megalocottus taeniopterus 26282
Cottus taeniopterus (non Kner, 1868) Myoxocephalus jaok 11366
Batrachocottus talievi Sideleva, 1999 Batrachocottus talievi 53210
Cottus tallapoosae Neely, Williams & Mayden, 2007 Cottus tallapoosae 72414
Radulinopsis taranetzi Yabe & Maruyama, 2001 Radulinopsis taranetzi 45347
Enophrys taurina Gilbert, 1914 Enophrys taurina 2850
Asemichthys taylori Gilbert, 1912 Asemichthys taylori 43939
Cottus tentaculatus Kner, 1868 Porocottus tentaculatus 559
Icelinus tenuis Gilbert, 1890 Icelinus tenuis 2850
Uranidea tenuis Evermann & Meek, 1898 Cottus tenuis 12965
Neocottus thermalis Sideleva, 2002 Neocottus thermalis 53210
Triglopsis thompsonii Girard, 1851 Myoxocephalus thompsonii 54621
Pseudoblennius totomius Jordan & Starks, 1904 Pseudoblennius totomius 41299
Ricuzenius toyamensis Matsubara & Iwai, 1951 Icelus toyamensis 40966
Cottus trachurus Pallas, 1814 Hemilepidotus hemilepidotus 93382
Cottus transsilvaniae Freyhof, Kottelat & Nolte, 2005 Cottus transsilvaniae 55856
Orthonopias triacis Starks & Mann, 1911 Orthonopias triacis 2850
Atopocottus tribranchius Bolin, 1936 Atopocottus tribranchius 559
Cottus tricuspis Reinhardt, 1830 Gymnocanthus tricuspis 12965
Gymnacanthus tricuspis orientalis Schmidt, 1927 Gymnocanthus tricuspis 114903
Cottus trigonocephalus Gratzianov, 1902 Leocottus kesslerii 57778
Myoxocephalus tuberculatus Soldatov & Pavlenko, 1922 Myoxocephalus tuberculatus 46206
Triglops uchidai Watanabe, 1958 Triglops scepticus 7297
Icelus uncinalis Gilbert & Burke, 1912 Icelus uncinalis 6876
Icelus uncinalis stenosomus Andriashev, 1937 Icelus stenosomus 27737
Cottus uncinatus Reinhardt, 1834 Artediellus uncinatus 58462
Gymnocanthus vandesandei Poll, 1949 Gymnocanthus vandesandei 46206
Cottus ventralis Cuvier, 1829 Gymnocanthus pistilliger 121848
Cottus verrucosus Bean, 1881 Myoxocephalus scorpius 43939
Myoxocephalus verrucosus ochotensis Schmidt, 1929 Myoxocephalus ochotensis 46206
Icelus vicinalis Gilbert, 1896 Icelus euryops 11366
Radulinus vinculus Bolin, 1950 Radulinus vinculus 2850
Abyssocottus werestschagini Taliev, 1935 Neocottus werestschagini 2058
Stlegicottus xenogrammus Bolin, 1936 Rastrinus scutiger 86856
Triglops xenostethus Gilbert, 1896 Triglops xenostethus 7297
Myoxocephalus yesoensis Snyder, 1911 Myoxocephalus yesoensis 46206
Bero zanclus Snyder, 1911 Alcichthys elongatus 43239
Argyrocottus zanderi Herzenstein, 1892 Argyrocottus zanderi 559
Hemilepidotus zapus Gilbert & Burke, 1912 Hemilepidotus zapus 2850
Pseudoblennius zonostigma Jordan & Starks, 1904 Pseudoblennius zonostigma 559
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cfm script by eagbayani, 10.5.01 ,  last modified by cmilitante, 29/11/12