| Citation | Plamoottil, M., 2014, Puntius nelsoni, Systomus chryseus and S. rufus (Cypriniformes: Cyprinidae), three new fish species from Kerala, India.. Int'l J of Fauna and Biological Stud. 1(6):135-145. |
| Ref. | 104747 Plamoottil, M., 2014 |
| Page | 145 |
| Named Used as Valid | Systomus sarana |
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